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After a chaotic encounter at Grand Central Station, the group is tranquilized and shipped toward a Kenyan wildlife preserve. However, a squad of militant penguins hijacks the ship, causing the four friends' crates to fall overboard and wash up on the shores of Madagascar. The second half of the film explores their struggle to survive in an environment where nature's rules—like the predator-prey relationship between Alex and Marty—begin to take over.